Infertile ova

[Trey Moore]

freshly emerged female H. euryalus in a trap in the evening. The next
morning she was paired with a male. They broke apart that evening , and the
male was released. I placed the female in a paper sack where she deposited
135 eggs over the next three days. after two weeks they have started to
collapse, and the ones I have opened had fluid in them but no developing
larvae.
   At about the same time I had a pairing of A. atlas, which stayed paired
for twenty four hours, the eggs are in the same condition as the above.
    What could the problem be?